In 2015, a research recovered mDNA from ancient canid specimens which were discovered on Zhokhov Island plus the Yana river, arctic Siberia. These specimens bundled the mandible of a 360,000–400,000 YBP Canis cf. variabilis (exactly where cf. is a Latin phrase meaning unsure). Phylogenetic analyses of those canids discovered nine mDNA haplotypes

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